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The idea of the gene has been a central organizing theme in contemporary biology, and the Human Genome project and biotechnological advances have put the gene in the media spotlight. In this book Lenny Moss reconstructs the history of the gene concept, placing it in the context of the perennial interplay between theories of preformationism and theories of epigenesis. He finds that there are not one, but two, fundamental – and fundamentally different – senses of ‘the gene’ in scientific use – one the heir to preformationism and the other the heir to epigenesis. ‘Gene-P’, the preformationist gene concept, serves as an instrumental predictor of phenotypic outcomes, while ‘Gene-D’, the gene of epigenesis, is a developmental resource that specifies possible amino acid sequences for proteins.